Tyrannidectes synallaxis

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Type material. Holotype male, paratypes 37 males and 61 females ex Synallaxis ruficapilla Vieillot, 1819 (Passeriformes: Furnariidae), BRAZIL, São Paulo State, Sete Barras, 24 ° 23 ' S 47 ° 55 ' W, 15 July 1979, Y. Oniki- Willis and E. O. Willis cols. (# Y- 27). Type deposition: Holotype male at DZUnesp-RC (# 3615); paratypes at DZUnesp-RC (# 3616 – 3668), MHNCI, ZISP, UMMZ, USNM, and DZSJRP. Male. (holotype, range for 8 paratypes in parentheses). Idiosoma, length × width, 342 (332 – 355) × 146 (130 – 145). Prodorsal shield: entire, lateral margins slightly concave at level of scapular setae, posterior margin sinuous, posterior angles acute, length 112 (111 – 118), width 100 (101 – 110), surface with a few rounded lacunae posterior to scapular setae (Fig. 7 A); bases of scapular setae se separated by 53 (50 – 53). Setae ve present, rudimentary. Scapular shields narrow. Humeral shields absent. Setae cp and c 2 situated on striated tegument. Subhumeral setae c 3 lanceolate, 28 (26 – 28) × 8 (7 – 9). Hysteronotal shield: greatest length 220 (215 – 228), width at anterior margin 95 (87 – 97), anterior margin sinuous, surface with small uniformly disposed lacunae. Distance between prodorsal and hysteronotal shields 10 (7 – 15). Posterior margins of opisthosomal lobes rounded. Terminal cleft shaped as an inverted U with divergent branches, 19 (18 – 23) long. Supranal concavity poorly distinct. Setae f 2 absent. Setae h 1 situated approximately at the same level as ps 3. Setae h 3 setiform, 23 (21 – 27) long; setae ps 2 83 (68 – 92) long; setae ps 1 filiform, about 5 long, situated slightly anterior to bases of setae h 3. Distances between dorsal setae: c 1: d 2 90 (85 – 92), d 2: h 1 95 (92 – 109), h 2: h 2 46 (43 – 49), h 3: h 3 30 (26 – 32). Epimerites I fused into a V, fused part tridentate posteriorly (Fig. 7 B). Coxal fields I, II without extensively sclerotized areas. Rudimentary sclerites rEpIIa absent. Coxal fields I – IV open. Epimerites IVa indiscernible. Genital arch 46 (43 – 49) in width; aedeagus 158 (148 – 156) long from anterior bend to tip, extending to tip of setae h 3. Genital papillae connected at bases. Genital and adanal shields absent. Adanal suckers 11 (10 – 13) in diameter, distance between centers of discs 26 (26 – 28), corolla indented, surrounding membrane with radial striae. Opisthoventral shields occupying lateral areas of opisthosoma; inner margins of these shields posterior to level of anal suckers with angular extensions, setae ps 3 inserted on anterior margins of these extensions. Setae 3 a and 4 b situated approximately at the same transverse level. Distance between ventral setae: 1 a: 4 b 103 (102 – 113), 4 b: 4 a 34 (33 – 38), 4 a: g 35 (38 – 45), g: ps 3 64 (62 – 68), ps 3: ps 3 56 (57 – 62). Femora I, II without crests (Fig. 9 A, B), other segments of legs I – IV without processes. Solenidion σ 1 of genu I 12 (10 – 14) long, situated at midlevel of segment; solenidion σ of genu III inserted at midlevel of segment. Genual setae cG I, II and mG I, II filiform. Seta d of tarsi II slightly shorter than corresponding seta f; seta d of tarsi III two times shorter than corresponding seta f (Fig. 9 C). Solenidion φ of tibia IV extending to midlevel of ambulacral disc. Tarsus IV 29 (24 – 30) long, without claw-like apical process; setae d and e button-like, seta d situated basally at level of seta r (Fig. 9 D). Female (range for 8 paratypes). Idiosoma, length × width, 485 – 518 × 162 – 175. Prodorsal shield: shaped as in male, length × width, 126 – 136 × 138 – 151, surface with small lacunae, bases of setae se separated by 71 – 76 (Fig. 8 A). Setae ve present, rudimentary. Scapular shields narrow, poorly developed dorsally. Humeral shields absent. Setae cp and c 2 situated on striated tegument. Setae c 3 lanceolate, 27 – 33 × 7 – 10. Anterior and lobar parts of hysteronotal shield separated, without integument between those shields (Fig. 8 A). Distance between prodorsal and anterior hysteronotal shields 16 – 30. Anterior hysteronotal shield roughly rectangular, anterior margin straight, greatest length 248 – 271, width at anterior margin 122 – 134, surface with numerous lacunae and a pair of lightly sclerotized areas at level of setae e 2. Length of lobar region 84 – 95, greatest width 84 – 96. Terminal cleft narrow, 68 – 94 long. Supranal concavity distinct; lobar shield undivided medially, surface without ornamentation. Setae f 2 absent. Setae h 1 at same level as supranal concavity. Setae h 2 lanceolate with blunt apex, 49 – 54 × 7 – 8. Setae ps 1 situated near inner margins of opisthosomal lobes, approximately equidistant between h 3 and h 2. Setae h 3 22 – 28 long. Distances between dorsal setae: c 1: d 2 99 – 120, d 2: h 1 139 – 156, h 2: h 2 67 – 75, h 3: h 3 30 – 48. Epimerites I fused into a V (Fig. 8 B). Coxal fields I – IV open. Epimerites IVa indiscernible. Translobar apodemes of opisthosomal lobes present, wide, fused to each other anterior to terminal cleft. Epigynum horseshoe shaped, greatest width 60 – 79; apodemes of ovipore connected with epimerites IIIa. Primary spermaduct with slight bulbous swelling near the head of spermatheca; secondary spermaducts 23 – 36 long (Fig. 17 H). Pseudanal setae filiform, setae ps 2 situated at level of posterior half of anal opening; distance between pseudanal setae: ps 2: ps 2 41 – 56, ps 3: ps 3 24 – 29, ps 2: ps 3 15 – 19. Femora II with ventral crest, other segments of legs I, II without processes. Solenidion σ 1 of genu I short, 14 – 18 long, situated at midlevel of segment. Solenidion σ of genu III inserted basally. Genual setae cG I, II, mG I, II as in male. Seta d of tarsi I, II slightly shorter than corresponding seta f, setae d of tarsi III, IV 2 – 3 times shorter than corresponding setae f. Genu IV dorsally inflated, with narrow dorsal crest.